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The rise and fall of testosterone levels initiates the hardening of the antler, the peeling off of antler velvet and casting or shedding of the antlers.
"The higher the testosterone level, the more susceptible the animal is to changes in temperature and other factors. If the animal does not have an adequate exposure to the cold and has more than a week's period of a cold spell, the antlers may soften," said David Kowalcyk, the director of the Institute of Vertebrate Zoology at the State University of New York, Stony Brook.
Kowalcyk, a renowned authority on carnivorans and the author of "The Zoo, the World and the Wild," added: "These are very powerful animals, animals that will have very long lives. How much influence will the testosterone exert on a short-lived animal? We don't know."
The researchers used the new technique to study the temperature of 20-year-old, adult adult male lions in the wild. The study was published online Oct. 14 in the journal Molecular Biology and Evolution.
Before the study, the researchers noted that although some studies had suggested that the testosterone levels may have effects on mating success, they had no idea how much of a change they might have made to the adult male behavior of this endangered species in the past four decades.
The lions' blood samples were collected from various body parts such as their eyes, ears, tail and hooves, then dried to preserve them. The researchers analyzed the blood to measure testosterone levels as well as cortisol levels, the hormone that tells the body to release stress hormones. It also measured the levels of testosterone and cortisol in the hair on the lions' horns and antlers.
To determine where the testosterone levels would affect the behavior of a male lion, the scientists measured the testosterone levels of the two highest and three lowest testosterone concentrations in his antlers in every animal tested over 20 years.
"We found that higher concentrations of testosterone occur during the winter, when the temperatures have been high for several weeks," Kowalcyk said. "Then, after the winter wears off, levels decrease for a few days at the beginning of summer season, and then increase again after the first of the summer months.
"After that, testosterone stays high for two weeks. This is where we expect to see an average decline in temperature as the animal grows old and matures."
The hormones released during puberty tell the body to prepare for the changes that occur as testosterone levels rise. For male lions, they can produce sperm earlier in puberty, with higher levels than adults, than
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